Abavorana decorata

Diagnosis. In addition to its phylogenetic placement (Fig. 1), Rana decorata is reassigned as a member of the genus Abavorana based on the combination of having a robust, medium-sized body; no vocal sacs in males; length of 1 st finger greater than 2 nd finger; disc width to finger width ratios of finger 3 and toe 4 is 1 - 1.5; dorsolateral fold indistinct or absent; the colour of the dorsolateral line being white or yellow; the humeral gland in males is prominent, raised and centrally positioned on the ventral surface of the upper arm; a weak or absent rictal ridge; outer metatarsal tubercle weak or absent; skin of dorsum smooth or finely shagreened; throat, abdomen and flanks smooth; posterior section of venter and back of the thigh rugose; no pale colouration on the margins of the tympanum; flanks dark-brown or black below the dorsal fold grading into a pale venter (Inger 1966; Oliver et al. 2015; Quah et al. 2017). Dorsum reddish-orange to rust-brown, encircled by a white or cream coloured dorsolateral line that encircles the snout, canthus rostralis, outer edge of the upper eyelids, and dorsum along the dorsolateral fold to the vent; lower flanks dark-brown or black below the dorsolateral line grading into a paler venter; dorsal colouration of the limbs light-grey or brown with speckling and prominent dark-brown or black bands. Abavorana decorata can be differentiated from its congeners on the basis of its ventral colour pattern which is reticulated in black and white especially on the lower flanks, underside of the limbs (especially hind-limbs) are boldly banded in black and white, and a prominent white streak under the eye and / or tympanum to the corner of the jaw (Figs 5, 6, 9 C-D). Adult males with SVL 46.21 - 56.0 mm, adult females with SVL 29.66 - 58.42 mm; adult males with SW 9.91 - 10.71 mm, adult females with SW 6.25 - 11.68 mm; adult males with IOD 5.59 - 5.95 mm, adult females with IOD 3.91 - 6.78 mm; adult males with TD 4.54 - 4.60 mm, adult females with TD 2.27 - 5.63 mm; adult males with FL 25.58 - 26.10 mm, adult females with FL 17.19 - 31.34 mm (Table 5). Distribution (Fig. 10). Endemic to Borneo. The species is known from Sabah state, Malaysian Borneo: Mount Kinabalu, Kampung Kiau Taburi, Danum Valley, Tawau Hills Park, Maliau Basin, Tambunan and Penampang (CalPhotos ID: 0000 0000 0912 1159); Sarawak state, Malaysia Borneo: Kubah National Park (Fig. 7 C) and Bario (Fig. 6 A-B); and West Kalimantan, Indonesia: Melawi Regency (Fig. 7 D). The species is expected to be wider ranging on the island. Figure 7. Living specimens of Abavorana decorata and Bornean A. luctuosa; A: Abavorana decorata from Tawau Hills, Sabah, East Malaysia. B: A. decorata from Maliau Basin, Sabah, East Malaysia. C: A. decorata from Kubah National Park, Sarawak, East Malaysia. D: A. decorata from Melawi Regency, West Kalimantan Province, Indonesia. E & F: A. luctuosa from Kubah National Park, Sarawak, East Malaysia. G: A. luctuosa (FMNH 238720) from Sipitang, Sabah, East Malaysia. H: A. luctuosa (FMNH 242883) from Sipitang, Sabah, East Malaysia. (Photographs by Alexander Haas [A, C & E], Paul Bertner [B], Muhammad Aini [D], Chien C. Lee [F], and Robert F. Inger [G & H], Figs G & H © Field Museum of Natural History. Created by Field Museum of Natural History, Amphibian and Reptile Collection and licensed under CC-BY-SA 4.0).