GBIF Taxonomic Backbone & Wikipedia Registry Profile
Description. Male (8.3 mm). Small size species. Vital body color is light-brown. Thoracic segments smooth dorsally (Fig. 16, 17 a, 28 d). Pleon segments 1 – 2 with small central tooth and 3 (rarely 2) lateral teeth on each side, submedial teeth is larger; pleon segment 3 with the posterior row of 7 teeth, central tooth small, marginal teeth are larger than the intermediate in the each lateral group of teeth. Urosome segment 1 with central tooth and with 2 lateral teeth on each side (1 large tooth and 1 small denticle); urosome 2 with 2 lateral pairs of acute teeth and with the stout cuspidate seta in the each pair. Head: (Fig. 17 b) subequal to pereonites 1 and 2 together, with an indistinct rostrum; eyes small suboval; anterior head lobe rounded; lower margin with a short notch. Antenna 1: (Fig. 17 c) nearly 84 % of body length, slender; flagellum exceeding 1.2 times the length of the peduncle, bearing 37 articles, few very thin and short setae scattered along flagellum; peduncle article 1 subegual to article 2 length, with 6 stout simple setae along posterior margin and with short acute protrusion at the distal posterior margin, article 3 0.39 times as long as article 2; accessory flagellum 6 - articulate. Antenna 2: (Fig. 17 d) flagellum almost 0.48 times as long as peduncle, 13 articles, with a crown of few thin setae on each article; peduncle articles 5 0.83 times as long as article 4, articles 1, 2, 3 together 1.3 times shorter than article 4; article 2 with protruding lobe reaching the distal end of article 3; articles 4 and 5 with numerous evenly spaced tufts of thin setae. Mouthparts. Labrum: (Fig. 17 e) frontal margin slightly concave medially, laterally pubescent. Mandible: (Fig. 17 f, g) incisor crenulate and stretching in a rounded tip; raker setae stout serrated, numerous (13 – 14), subequal to incisor in length; molar flattened triturative; palp 3 - articulate, palp segment 2 with 6 – 7 rare setae along posterior margin, palp segment 3 with 9 – 11 setae, distal setae are larger, 1 - st article with protruding lobe, the percentage of total length of the articles of the palp 18 % (article 1), 35 – 41 % (article 2) and 41 – 47 % (article 3). Labium: (Fig. 17 h) inner lobes small but well defined, rounded; outer lobes rounded, with minute setae at tips. Maxilla 1: (Fig. 17 i) inner plate triangulate, lined with a row of 11 setae; outer plate elongate, 1.7 times as long as inner plate, apically with 2 rows of 9 bifurcate strong setae; palp 2 - articulate, article 2 2.2 times as long as article 1, article 2 apical margin with two rows of 9 – 10 simple setae each. Maxilla 2: (Fig. 17 j) inner plate slightly shorter than outer plate, facial setae of inner plate numerous, located submarginally, tip covered with numerous simple setae; outer plate with numerous simple setae on the tip. Maxilliped: (Fig. 18 a) inner plates medium size, fully cleft, distal margin with a row of 3 teeth, inner margin with long plumose setae (9 – 11 setae); outer plates reaching 4 / 5 length of palp article 2, inner margin lined with a row of short stout teethlike setae longer toward the apex, smoothly into the apical row of long thin setae, with several transverse bundles of short setae submarginally; palp 4 - articulate, the percentage of total length of the articles of the palp are 14 % (article 1), 33 % (article 2), 25 % (article 3), 28 (article 4), article 2 with numerouse setae along inner margine; tip of article 3 with a crown of long simple setae; article 4 inner margin with a line of simple setae. Pereopod 1 (gnathopod 1): (Fig. 18 b, c) coxa extended distally, with rounded anterior corner; basis straight with a row of short simple setae along anterior margin and with rare long setae along posterior margin; ischium with dense tuft of long setae at the postero-distal corner; merus subrectangular, with a crown of simple setae along distal margin and with a long spot of dense short setae at the posterior margin; carpus 1.2 times as long as propodus, 8 – 9 oblique rows of long setae along outer posterior margin, with a row of long setae along anterior part of distal margin and tomentose at the anterior-distal corner; propodus suboval, with few groups of long setae along posterior margin, with 5 tufts of long simple setae along anterior margin and with a dense bunch of long simple setae at the anterior distal corner, palm oblique rounded, smoothly passes in posterior margin, minutely serrate with a row of numerous short stout simple setae, outer surface with a transverse row of long setae distally; dactylus as long as palm, curved near the tip, with nail and with short tooth-like protrusion above it. Pereopod 2 (gnathopod 2): (Fig. 18 d, e) coxa extended, rounded distally, with acute teeth at the posterior distal corner; gills with indistinct peduncle; basis slightly expands medially, as long as coxa, with a row of short setae along anterior margin and with a few long simple setae along posterior margin; merus subrectangular with long acute protrusion at the posterior distal corner, without setae along posterior margin; carpus triangular, 4 times shorter than propodus; propodus extends distally, palm oblique, with 3 slight protuberances at the proximal half, palmar corner with large acute tooth, posterior margin with numerous tufts of setae, inner surface with a fosse and two short stout setae near the palmar corner; dactylus slightly inflated in the middle part, slightly shorter than palm, slightly curved to the inside of propodus, outer margin with a few short setae, tip of dactylus obtuse. Pereopod 3: (Fig. 19 a) coxa long, subequal to coxa 2, with parallel anterior and posterior margins, rounded distally; basis linear, slightly curved in the proximal 1 / 5, with rare minute setae along anterior and posterior margins; merus long, with 3 groups of strong simple setae along anterior margin and with a single cluster of simple setae at the anterior distal corner, posterior margin with 4 tufts of simple setae, posterior distal corner with a bunch of simple setae; carpus 0.79 times as long as merus, with 6 groups of strong simple setae along posterior margin; propodus linear, as long as carpus, with 6 groups of short strong simple setae along posterior margin; dactylus small (0.3 times length of propodus), with nail. Pereopod 4: (Fig. 19 b) coxa shorter than coxa 3, anterior and posterior corners rounded, posteriorly excavated; segements 3 – 7 shorter than in pereopod 3; merus with two groups of strong simple setae along anterior margin; carpus with 5 groups of strong simple setae along posterior margin; propodus with 6 groups of short strong simple setae along posterior margin; dactylus medium (0.36 times length of propodus), with nail. Pereopods 5 – 7 elongated from pereopod 5 to pereopod 6, pereopods 6 and 7 equal in length. Pereopod 5: (Fig. 19 c, d) coxa with rounded front margin, with sloped hind margin, with rounded posterior distal corner and with rounded anterior lobe pulled down; basis regular, with narrow posterior wing, width 0.61 times as long, posterior distal corner rounded, hind margin crenulated, anterior margin with small stout setae; merus slightly broaded medially, with 3 short stout setae along anterior margin, with 2 short stout setae at the middle of posterior margin, with a cluster of stout setae at anterior and posterior distal corners each; carpus 0.93 times as long as merus, with 2 groups of 2 strong setae at the anterior margin and with a single cluster of strong stout setae at the anterior and posterior distal corners each; propodus linear, more narrow and 1.45 times as long as carpus, with 6 bunches of strong simple setae along anterior margin; dactylus medium (0.36 times length of propodus), with nail. Pereopod 6: (Fig. 19 e, f) coxa with sloped hind margin, posterior distal corner distinct, with acute anterior lobe pulled back and down, front margin with few short simple setae; basis regular, slightly narrowed distally, with posterior wing, width 0.59 times as long, posterior distal corner slightly rounded, hind margin crenulated, anterior margin with small stout setae; merus long, with 3 – 4 short stout simple setae along anterior and posterior margins each, and with cluster of strong stout setae at anterior and posterior distal corners each; carpus 0.88 times as long as merus, with 2 groups of 2 stout setae at the anterior margin and with a single cluster of strong stout setae at the anterior and posterior distal corners each; propodus linear, more narrow and 1.4 times as long as carpus, with 6 clusters of short cuspidate setae along anterior margin; dactylus medium (0.36 times length of propodus), with nail. Pereopod 7: (Fig. 19 g, h) coxa small, without distinct anterior lobe and with rounded posterior lobe; basis regular, with rounded posterior wing, width 0.64 times of length, posterior distal corner rounded, posterior margin crenulated, anterior margin with small stout setae; merus long, with 3 groups of stout setae along anterior margin, with two tufts of stout setae along posterior margin, and with single cluster of strong setae at anterior and posterior distal corners each; carpus, propodus and dactylus similar to that in pereopod 6. Coxal gills 2 – 6. Coxal gill 6 slightly smaller than gill 5. Epimeral plates: (Fig. 20 a, b, c) plate 1 with oblique crest, posterior margin slightly rounded, with 1 unclear notch, lower margin without armaments, hind corner subquadrate; plate 2 posterior margin slightly concave, ventral margins convex, with oblique crest, posterior margin with 2 indistinct notches, ventral margin without armaments; plate 3 hind corner produced, acute, posterior margin concave, with 3 acute teeth, ventral margin with a row of 5 – 6 strong simple setae. Pleopods: (Fig. 20 d – h) normal; peduncle with two specific toothed coupling setae on the inner corner; coupling basis of inner ramus with 3 specific stout two-pointed cuspidate setae along inner margin. Uropod 1: (Fig. 20 i) peduncle with strong interramal spur; rami subequal in length, 0.9 – 0.91 as long as peduncle; peduncle and rami with tiny simple setae along margins, both rami with few tiny simple setae at the tip. Uropod 2: (Fig. 20 j) peduncle shorter than outer ramus (0.76 ramus length), with acute protrusion at the distal margin; inner ramus shorter than outer, with numerous simple tiny setae along outer margin; uropod 2 shorter than uropod 1. Uropod 3 is lost. Telson: (Fig. 20 k) fully cleft; lobes diverging distally, proximal notch positioned medially; apical and subapical simple cuspidate setae long, lobes with single short cuspidate setae at the medial margin each. Females unknown.
