GBIF Taxonomic Backbone & Wikipedia Registry Profile
Description of the holotype. Adult female (MUTPL 932; Figs 2, 3, 5 A), medium sized, with many large and small yellowish white eggs. Head slightly narrower than the body, wider than long, head length 91 % of head width, head width 34 % of SVL, head length 31 % of SVL; snout relatively short, snout to eye distance 13 % of SVL, eye-nostril distance 24 % of head length; snout rounded in dorsal view, sloping in profile; nostrils slightly elevated producing a shallow depression in the internarial area; canthus rostralis indistinct in dorsal view, rounded in cross section; loreal region slightly concave; lips non-flared; moderate sized eyes (sensu Guayasamin et al. 2020), eye diameter 11 % of SVL, 34 % of head length; eye-nostril distance 69 % of eye diameter; eye diameter 93 % of IOD; eyes directed anterolaterally at ~ 40 ° from midline, slightly visible from below; upper eyelid width 85 % of IOD; tympanic annulus and tympanic membrane evident, but the membrane with coloration similar to that of surrounding skin; tympanum oriented slightly dorsolaterally; tympanum large (sensu Guayasamin et al. 2020), its diameter ~ 46 % of eye diameter; weak supratympanic fold present, slightly concealing the upper margin of the tympanum; choanae large, ovoid, not concealed by palatal shelf of maxillary arch, closer to the distal margin of the dentigerous processes of vomers than to the margin of mouth; dentigerous processes of vomers ovoid, in transverse row between the choanae, separated medially by distance slightly lower than the width of processes; each process bearing three teeth; tongue just slightly longer as wide, not notched posteriorly, and only ~ 1 / 5 of posterior part not adherent to the floor of mouth. Skin of dorsal surfaces shagreen with elevated, and some enameled, warts corresponding to white spots; throat smooth; ventral skin coarsely areolate; ventral surfaces of thighs below vent with a pair of large, round, flat tubercles (subcloacal warts); cloacal opening directed posteriorly at upper level of thighs, no distinct cloacal sheath; cloacal region bordered ventrally by many enameled (white) warts. Upper arm thin, forearm somewhat robust; outer edge of forearms with row of enameled warts that continue into the external edges of Finger IV; hand length 34 % of SVL; palmar tubercle large, elliptical; thenar tubercle large, ovoid; subarticular tubercles prominent, round and round in section; numerous round palmar supernumerary tubercles present, much smaller than subarticular tubercles; relative lengths of fingers I <II <IV <III; concealed prepollex; fingers with broad lateral fringes; webbing absent between Fingers I and II, basal between II and III, moderate between outer fingers: III 2 + - 2 IV (Fig. 5 A); bulla absent; discs on fingers expanded, truncate; disc pads nearly triangular. Hindlimbs long, slender; femur length 55 % of SVL; tibia length 57 % of SVL; foot length 48 % of SVL; outer edge of tarsus with row of enameled warts that often continue into the external edges of Toe V; inner edge of tarsus bearing a long fold; inner metatarsal tubercle large, elliptical; outer metatarsal tubercle indistinct; subarticular tubercles rounded and flat; plantar supernumerary tubercles inconspicuous; relative length of toes I <II <III <V <IV; toes with broad lateral fringes; webbing between toes moderate: I 1 --- 2 II 1 -- 2 III 1 -- 2 IV 2 - 1 + V (Fig. 5 A); discs on toes expanded, truncate, lacking papillae; disc pads nearly triangular.
