Hemiphractus kaylockae

Description of the holotype. AN ADULT FEMALE: SVL 54.2 MM (NOT INCLUDINg PROBOSCIS); TIBIA LENgTh 28.4 MM (NOT INCLUDINg CALCAR); FOOT LENgTh 25.3 MM; hEAD LENgTh 24.3 MM (NOT INCLUDINg PROBOSCIS); hEAD wIDTh 26.5 MM; SNOUT-PARAOCCIPITAL PROCESS LENgTh 25.1 MM; TYMPANUM OVOID, 6.6 MM hIgh X 3.6 MM wIDE; EYE DIAMETER 6.5 MM; INTERORBITAL DISTANCE 10.2 MM; EYE – TYMPANUM DISTANCE 5.4 MM; BODY MODERATE; hEAD LARgE, wIDER ThAN BODY, gREATEST wIDTh AT LEVEL OF PARS ARTICULARIS, BEARINg PROMINENT PARAOCCIPITAL PROCESSES; PARAOCCIPITAL PROCESSES PROjECTINg POSTEROLATERALLY IN DORSAL VIEw, EXTENDINg POSTERIOR TO LEVEL OF PARS ARTICULARIS; SkIN CO-OSSIFIED OVER DORSAL SURFACE OF SkULL; OCCIPITAL RIDgE PRESENT, wELL-DEVELOPED; PARAOCCIPITAL RIDgES PRESENT, wEAkLY DEVELOPED; SUPRAORBITAL, zYgOMATIC, PREORBITAL, AND CANThAL RIDgES ABSENT; CANThUS ROSTRALIS INDISTINCT; LOREAL REgION FLAT; PROBOSCIS ShORT, FLEShY, POINTED, LATERALLY FLATTENED IN CROSS SECTION; SNOUT ANgLINg POSTERIORLY AT LESS ThAN 45 o FROM hORIzONTAL; NOSTRILS OVOID, SLIghTLY PROTUBERANT, PROjECTINg DORSALLY; LIPS NOT FLARED. TOP OF hEAD RELATIVELY FLAT; INTERORBITAL REgION 38 % OF hEAD wIDTh; EYE DIAMETER 25 % hEAD wIDTh; TYMPANUM VERTICALLY OVOID, ORIENTED POSTERIORLY AND SLIghTLY LATERALLY, VISIBLE BELOw PARAOCCIPITAL PROCESSES IN DORSAL VIEw, gREATEST DIAMETER EqUIVALENT TO EYE DIAMETER; wIDTh OF TYMPANUM 1.6 TIMES EYE – TYMPANUM DISTANCE; UPPER EYELID wITh ONE ENLARgED POINTED PROjECTION AND SCATTERED SMALLER TUBERCLES; INTERORBITAL TUBERCLE ABSENT; POSTORBITAL TUBERCLE ABSENT; SkIN OVER POSTERIOR SURFACE OF PARS ARTICULARIS BEARINg ONE SMALL, FLEShY TUBERCLE. TONgUE LARgE, OVOID; VOCAL SLITS ABSENT; MANDIBLE BEARINg SERRATIONS ALONg ENTIRE LENgTh OF LOwER jAw, wITh TwO PROMINENT, TUSk-LIkE ODONTOIDS PROjECTINg FROM MENTOMECkELIAN BONES. AXILLARY MEMBRANE ABSENT; ThORACIC FOLD ABSENT; DERMAL FOLDS ON wRISTS ABSENT; FINgERS LONg, SLENDER, BEARINg ENLARgED DISTAL PADS; FINgER PADS ABOUT 1.8 TIMES wIDTh OF DIgITS AT PENULTIMATE PhALANgE; RELATIVE LENgThS OF FINgERS: III> IV> II> I; LATERAL FRINgE ON FINgERS ABSENT; OUTER METACARPAL TUBERCLE LOw, OVOID; INNER METACARPAL TUBERCLE OVOID, DIFFUSE; FOREARM wITh SCATTERED TUBERCLES, LARgE, POINTED, VARIOUSLY FUSED TO PRODUCE AN EVIDENT LATERAL FRINgE; wEBBINg ON hANDS ABSENT. HEELS OF ADPRESSED hIND LIMBS OVERLAP; TIBIOTARSAL ARTICULATION EXTENDINg TO SNOUT whEN hIND LIMB EXTENDED; TARSAL FOLD PRESENT, wEAkLY DEVELOPED; TIBIA LENgTh 52 % SVL; FOOT LENgTh 47 % SVL; OUTER METATARSAL TUBERCLE ABSENT; INNER METATARSAL TUBERCLE PRESENT, LARgE, OVOID; TOES IV AND V wITh LATERAL FRINgE, PROMINENT ON TOE V; FRINgE OF TOE V EXTENDINg ONTO OUTER SURFACE OF BASE OF TARSUS; DORSAL SURFACE OF hIND LIMB RELATIVELY SMOOTh, BEARINg IRREgULAR SETS OF TUBERCLES ALONg ThREE TRANSVERSE BANDS ON ShANk; CALCAR PRESENT, SMALL; RELATIVE LENgThS OF TOES: IV> V> III> II> I; TOE PADS ABOUT 1.5 TIMES wIDTh OF PENULTIMATE PhALANgE; wEBBINg BASAL. CLOACAL OPENINg DIRECTED POSTEROVENTRALLY AT MIDLEVEL OF ThIgh. DORSUM SMOOTh wITh SCATTERED, LOw, DIFFUSE TUBERCLES; FLANkS SMOOTh; VENTRAL SURFACES AND ThROAT SMOOTh wITh SCATTERED LOw, DIFFUSE TUBERCLES; SACRAL DIAPOPhYSES PROjECTINg LATERALLY, SLIghTLY NARROwER ThAN POSTERIORMOST TRANSVERSE PROCESSES OF ALL VERTEBRAE; LATERAL TERMINI OF TRANSVERSE PROCESSES OF PRESACRAL VERTEBRAE VI – VIII NOT CO-OSSIFIED wITh SkIN; FIVE NEURAL SPINES PROjECTINg DORSALLY, CO-OSSIFIED wITh SkIN, VISIBLE EXTERNALLY. PUPIL hORIzONTAL. ALL DORSAL SURFACES UNIFORMLY BROwN (FIg. 4). ThIgh BEARINg TwO, ShANk BEARINg ThREE INDISTINCT TRANSVERSE BANDS OF IRREgULAR, BROwN MOTTLINg ON DORSAL SURFACES, CORRESPONDINg wITh ThREE BANDS OF TUBERCLES ON ThESE BANDS; TOES I AND II ARE UNIFORMLY BEIgE, TOES III – V UNIFORMLY BROwN; FOREARM DARk BROwN, BEARINg BROAD, BROwN, TRANSVERSE BAND ON DORSAL SURFACE; DORSAL SURFACE OF MANUS MORE PALE ThAN FOREARM, ULTIMATE AND PENULTIMATE PhALANgES OF FINgERS I AND II BEIgE, FINgERS III AND IV DARk BROwN. UPPER AND LOwER LIPS wITh NARROw, TRANSVERSE, DARk BROwN BANDS, NOT EXTENDINg ONTO PROXIMAL SURFACES. FLANkS BEIgE wITh DIFFUSE BROwN PUNCTATIONS; VENTER BEIgE wITh SCATTERED DIFFUSE, FINE BROwN MARkINgS; ThROAT DARk BROwN, EXTENDINg ONTO PECTORAL REgION. VENTRAL SURFACES OF FOREARMS BEIgE wITh FINE, BROwN MOTTLINg; VENTRAL SURFACES OF hANDS BEIgE. VENTRAL SURFACES OF hIND LIMB AND FOOT BEIgE wITh FINE, PALE BROwN MOTTLINg. Variation. CONSISTENT SEXUAL DIMORPhISM IN COLORATION OR SkIN TEXTURE wAS NOT EVIDENT (FIgS. 4, 8). GRANULATION ON ThE ThROAT IN Hemiphractus kaylockae IS VARIABLE AMONg SPECIMENS EXAMINED, VARYINg FROM SMOOTh (E. g., KU 93506, 93507, 107411, 107417, 154992) TO SMOOTh wITh TUBERCLES (E. g., AMNH 92668, KU 107419), SMOOThLY gRANULAR wITh NO TUBERCLES (E. g., KU 107412, 107413), TO SMOOThLY gRANULAR wITh TUBERCLES PRESENT (E. g., KU 107408, 107414, 107422, 107423, 154991, UMMZ 152624). GRANULATION OF ThE VENTER RANgES FROM SMOOTh TO UNIFORMLY gRANULAR (E. g., KU, 107411, 107422, 154992, UMMZ 152624); SOME SPECIMENS wITh SMOOTh OR UNIFORMLY gRANULAR VENTERS ALSO hAD VARIABLE NUMBERS OF DISCRETE TUBERCLES PRESENT (E. g., AMNH 92668, KU 93506, 93507, 107412, 107413, 107414, 107417, 107419, 107423, 154991). ThROAT COLORATION IN MOST SPECIMENS IS TAN (E. g. KU 107417) TO BROwN OR DARk BROwN (E. g., AMNH 92668, KU 93506, 107408, 107411, 107412, 107419, 107422, 154992) wITh SOME ShOwINg PROMINENT whITE MOTTLINg, STRIPINg, AND / OR PROMINENT LABIAL BARRINg (E. g. KU 93506, 107412, 107422, 154992) AND KU 93507, 107413, 107414, 107417, 107423, 154991, UMMZ 152624 DISPLAYINg STIPPLINg OR SPECkLINg OF BROwN AND whITE. VENTRAL COLORATION IN MOST SPECIMENS TYPICALLY IS A CONTINUATION OF ThE gULAR COLORATION OF DARk BROwN OR TAN EXTENDINg TO APPROXIMATELY MID-ABDOMEN whERE IT ABRUPTLY ChANgES TO A BEIgE COLORATION ThAT TYPICALLY CONTAINS VARIABLE AMOUNTS OF DARkER BROwN STIPPLINg (E. g., AMNH 92668, KU 93507, 107408, 107411, 107414, 107417, 107419, 107423, 154992); KU 93506 AND KU 107413 ShOw A SIMILAR PATTERN hOwEVER ALSO DISPLAY whITE MOTTLINg IN ThE PECTORAL REgION whILE KU 107422 DISPLAYS whITE SPOTTINg POSTERIORLY. ThE DORSUM OF H. kaylockae IS SMOOTh OR UNIFORMLY SMOOThLY gRANULAR IN ALL SPECIMENS wITh KU 93506, 107423, 154991, 154992, AND UMMZ 152624 hAVINg SOME TUBERCLES. DORSAL COLORATION IN MOST IS AN OVERALL TAN OR BEIgE ThAT MAY OR MAY NOT INCLUDE SOME LEVEL OF DARkER BROwN STIPPLINg OR MOTTLINg (E. g. KU 107408, 107412, 107413, 107417, 107419, 107422, 107423, 154991, 154992) whILE SOME SPECIMENS DISPLAY AN OVERALL UNIFORM BROwN COLOR (E. g. AMNH 92668, KU 93506, 93507, 107411, 107414). SOME SPECIMENS ShOw A PROMINENT DARk BAND ON ThE FOREARM (E. g. KU 107408, 107412, 107422, 107423, 154992). HOwEVER, ThIS BAND IS DIFFUSE OR wEAkLY DEVELOPED IN ALL OThER SPECIMENS. TwO OR ThREE DARk TRANSVERSE BANDS ARE PRESENT ON ThE FEMORAL AND TIBIAL REgIONS (ABSENT OR BARELY DISCERNIBLE ON AMNH 92668, KU 93506, 107411, 107419, 154991), AND A DIFFUSE OR VERY EVIDENT DARk BAND OR ChEVRON PRESENT ON ThE SACRAL REgION (E. g., KU 107408 [wEAkLY DEVELOPED], 107412, 107414, 107417, 107422, 154992, UMMZ 152624); IN SPECIMENS KU 107422 AND 154992 ThERE ARE TwO ChEVRONS PRESENT; IN ADDITION, SPECIMENS KU 107414, 107417, 107422, AND 154992 hAVE A DARk IRREgULAR VERTEBRAL STRIPE EXTENDINg FROM ThE SACRAL BAND CONNECTINg TO A DARk BLOTCh AT ThE OCCIPUT. ThE POSTERIOR SURFACE OF ThE ThIgh OF KU 107412 – 14 AND UMMZ 152624 BEARS A PATTERN OF TAN OR BROwN MOTTLED wITh whITE OR BEIgE whILE ALL OThER SPECIMENS BEAR A DIFFUSE (E. g., KU 93506, 93507, 107411, 107422, 107423, 154991, 154992) OR DISTINCT whITE STRIPE ALONg ThIS SURFACE (E. g., AMNH 92668, KU 107408, 107417, 107419).