Miconia kellyana

Diagnosis: — Miconia kellyana can be distinguished from M. chinantlana by its upper internodes that are covered with a mixture of simple trichomes, apically bifid trichomes, and stalked-stellate trichomes (vs. a mixture of only short-stalked and long-stalked stellate trichomes), petioles 2.5 – 7 mm long (vs. 13 – 48 mm long), elliptic-lanceolate 5 - plinerved leaf blades that are 1.4 – 3.5 cm wide, basally obtuse to cuneate and never oblique (vs. ovate to lance-ovate 5 – 7 - plinerved blades that are 4 – 9.5 cm wide, basally broadly rounded to subcordate and oblique on some leaves), and hypanthia copiously covered with simple spreading trichomes intermixed with some apically bifid trichomes and a few stalked-stellate trichomes (all 1 – 1.5 mm long) that completely conceal the surface at anthesis (vs. covered with spreading stalked stellate trichomes mostly 0.25 mm long and not concealing the surface at anthesis). Shrubs 2 – 3 m tall, cauline internodes of uppermost branchlets rounded, copiously covered with a mixture of brown simple trichomes, stalked-stellate trichomes and apically bifid trichomes 0.5 – 1 mm long becoming glabrate with age. Leaves of a pair equal to unequal in size, concolored when dry, petioles 2.5 – 7 mm long, canaliculated adaxially and densely covered with an indumentum like that of upper internodes; blades 5 – 13.5 × 1.4 – 3.5 cm, membranaceous when dry, lanceolate to elliptic-lanceolate, apex attenuate, base obtuse to cuneate, margin essentially entire, 5 - plinerved with an elevated primary midvein and one pair of secondaries mostly alternately diverging from the midvein 0.9 – 2.5 cm above the blade base, the tertiary veins also conspicuous, elevated and spaced 3 – 4 mm apart at the wider portions of the blade, adaxially copiously beset with ± appressed smooth trichomes 0.25 – 1 mm long, abaxially prevailingly covered with spreading stalked-stellate trichomes 0.5 1 mm long intermixed with some apically bifid and simple trichomes of the same length on primary, secondary and higher-order veins, and on the blade surface. Inflorescence terminal, thrysoid, 3 – 5 cm long including a peduncle 0.5 – 1 cm long, copiously covered with spreading yellowish smooth apically bifid trichomes mostly 0.5 – 1 mm long; bracts of the rachis nodes 1.5 – 3 mm long, sessile, persistent, subulate, glabrous adaxially, abaxially sparsely covered with smooth and / or apically bifid trichomes 0.25 – 0.5 mm long; bracteoles 1.5 – 2 mm long, sessile, persistent, linear-subulate, indumentum on the abaxial surface like that of the bracts. Flowers 5 (6) - merous, perfect, subsessile or on pedicels 0.5 mm long, hypanthia campanulate, 3 – 4 × 3 mm, copiously covered with yellowish smooth simple and some inconspicuously apically bifid trichomes 1 – 1.5 mm long that completely conceal the surface at anthesis; calyx tube 0.5 mm long, the calyx lobes 1 × 1.5 mm, bluntly triangular (at anthesis), glabrous adaxially and covered with indumentum like that of the hypanthia and calyx teeth abaxially, the exterior calyx teeth 5 (6), subulate, 4 – 6 mm long and ca. 0.12 mm or less wide at the base, exceeding the calyx lobes in length and concealing them; torus glabrous adaxially. Petals 3 – 4 × 2.5 mm, white becoming pinkish with age, glabrous, obovate, apically rounded, apex rounded and beset with gland-tipped trichomes. Stamens 10 (12), isometric and isomorphic, forming a ring around the style at anthesis but becoming deflexed to one side of the flower opposing the style, filaments 2 – 2.5 mm long, glabrous, complanate, subulate, white becoming pinkish-red with age; anthers 3 – 4 × 0.5 mm, yellow but turning reddish-pink with age, oblong, broadly rounded to ± truncate at the apex with a ventrally inclined apical pore; connective slightly thickened dorsally and prolonged ventro-basally above the filament insertion into a bilobulate ventral appendage less than 0.25 mm long. Ovary (at anthesis) ca. 2 / 3 inferior, 5 - locular, ± globose, apex rounded and beset with gland-tipped trichomes. Styles straight and curved distally but typically becoming declinate, 6 – 8 mm long, white, sparsely to moderately beset with gland-tipped trichomes up to 0.25 mm long for the basal ⅓ to ½ of their length; stigmas punctiform. Mature berries and seeds not seen. Paratypes: ― HONDURAS. Cortés: Parque Nacional El Cusuco (CNP), Sierra del Merendón, W of San Pedro Sula. Plot GU / 2 / SS 5, 2000 m by trail GU 2, SE from Guanales campsite, [ca. 15.48020 ° N, 88.22665 ° W], 1505 m, 15 June 2013, D. L. Kelly & A. C. Dietzsch 15016 (CAS!, EAP!); CNP, 1100 m by trail GU 1 from Guanales campsite, 15.49420 ° N, 88.23832 ° W, 1600 m, 2 August 2018, A. Ward CUGU 2018 01 (NY); CNP, Guanales trail GU 1, 15.49486 ° N, 88.23832 ° W, ca. 1650 m, 2 August 2018, A. Ward CUGU 2018 02 (BM, CAS!); CNP, 2000 m by trail SE from Guanales campsite, [ca. 15.48020 ° N, 88.22665 ° W], 1505 m, 2 August 2018, A. Woolf CUGU 2018 03 (CAS!, MO); CNP, near Guanales campsite, 15.48124 ° N, 88.22294 ° W, 1532 m, 23 June 2019, A. Ward CUGU 2019 01 (CAS!, EAP); CNP, by trail GU 2, SE from Guanales campsite, [ca. 15.494403 ° N, 88.237927 ° W], 1575 m, 23 June 2019, A. Ward CUGU 2019 03 (P); CNP, by trail GU 2, SE of Guanales campsite, [ca. 15.493993 ° N, 88.237964 ° W], ca. 1550 m, 23 June 2019, A. Ward CUGU 2019 02 / 04 (HEH).