Nidirana guangdongensis

Description of holotype. SYS a 005767 (Figs 5, 6), adult male. Body large and elongated, SVL 55.2 mm; head longer than wide (HDW / HDL 0.90), flat above; snout rounded in dorsal and lateral views, slightly protruding beyond lower jaw, longer than horizontal diameter of eye (SNT / ED 1.30); canthus rostralis distinct, loreal region concave; nostril round, directed laterally, closer to the snout than to the eye; a longitudinal swollen mandibular ridge extending from below nostril through lower edges of eye and tympanum to above insertion of arm, where the ridge is intermittent, forming a maxillary gland and shoulder gland; supratympanic fold absent; interorbital space flat, narrower than internasal distance (IND / IOD 1.24); pupil elliptical, horizontal; tympanum distinct, round, TD / ED 0.86, and close to eye, TED / TD 0.32; pineal ocellus slightly visible; vomerine ridge present, bearing small teeth; tongue large, cordiform, notched behind; a pair of subgular vocal sacs present. Forelimbs moderately robust, lower arm 0.17 of SVL and hand 0.27 of SVL; fingers thin, relative finger lengths II <I <IV <III; tip of each finger slightly dilated, forming rounded disks; lateroventral grooves on all fingers except finger I, not meeting at the tip of disks; fingers free of webbing; presence of distinct lateral fringes on inner and outer sides of fingers II, III and IV, and on outer side of finger I; subarticular tubercles prominent and rounded; week supernumerary tubercles below the base of each finger; three elliptic, large, prominent and very distinct palmar tubercles; a single nuptial pad on the dorsal surface of first finger, nuptial spinules invisible. Hindlimbs relatively robust, tibia 0.54 of SVL and foot 0.77 of SVL; heels overlapping when hindlimbs flexed at right angles to axis of body; tibio-tarsal articulation reaching the nostril when hindlimb is stretched along the side of the body; toes relatively long and thin, relative lengths I <II <V <III <IV; tip of each toe slightly dilated with remarkable elongated ventral callous pad, forming long and pointed disk; well-developed lateroventral grooves on toes, not meeting at the tip of disks; webbing moderate, webbing formula: I 1 ⅓ - 2 II 1 ⅓ - 2 ⅓ III 1 ⅔ - 3 IV 3 ⅓ - 1 ⅓ V; presence of lateral fringes on inner and outer sides of each toes, forming distinct dermal flap on the lateral edges of toes I and V; subarticular tubercles rounded, prominent; inner metatarsal tubercle elliptic, length triple the width; outer metatarsal tubercle indistinct, small and rounded; tarsal folds and tarsal tubercle absent. Dorsal surface rough with dense horny spinules; developed dorsolateral fold with sparse horny spinules from posterior margin of upper eyelid to above groin but intermittent posteriorly; flank rough with dense tubercles and dense horny spinules; a large and smooth suprabrachial gland behind base of forelimb, prominent; dorsal surface of forelimb relatively smooth without horny spinules, weak longitudinal ridges on upper arms and slightly extending to lower arm; the dorsal surfaces of thigh and tibia rough with dese tubercles and dense horny spinules, forming several longitudinal ridges. Ventral surface of throat, body, and limbs smooth; large flattened tubercles densely arranged on the rear of thigh and around vent.